- Chemical ecology of arthropods.
- Ecology of the hard tick Ixodes ricinus L. and its role as a vector of human diseases.
- Interactions between ticks, environment and hosts.
- Host-parasite interactions: changes in physiology and manipulation of tick behavior by associated pathogenic microorganisms.
- Strategies to reduce the risks of tick-borne diseases transmission.
- Beekeeping for rural development.
- Honeybee Apis mellifera L. parasites and pathogens.